Selected material studied: Caravaca de la Cruz, Sierra de la Zarza, 30SWG6798, 1285m, 2-V-2002, Cano 836b (MUB 13075).
Basic soils and base of trunk of Quercus rotundifolia.
Previous records: Ros & Guerra (1987) sub Tortula ruraliformis var. subpapillosissima; Gallego(2002); Gallego et al. (2002a); Cano et al. (2004); Guerra et al. (2006).